Rhipsalis cereuscula Haworth 1830

Hariota cereuscula (Haworth) Kuntze 1891, Erythrorhipsalis cereuscula (Haworth) Volgin 1981 Hariota sagllonis Lemaire 1838, Rhipsalis saglionis (Lemaire) Otto 1843

Rhipsalis brachiata W. J. Hooker 1843 Rhipsalispenduliflora N. E. Brown 1877 Rhipsalis simmleri Beauverd 1907

Plants shrubby to bushy, to 60 cm (24 in) long, much branched, usually pendent, growth determinate, with composite areoles. Stem segments strongly dimorphic; long shoots cylindrical, almost round in cross section, 10-30 cm

(3.9-12 in) long, 3-4 mm in diameter; short shoots borne in apical clusters, branching widely apically, obscurely four- or five-angled, 1-3 cm (0.4-1.2 in) long, 3-4 mm (to 0.2 in) in diameter. Areoles marginal, with two to four short bristles. Flowers borne apically in spring, bell shaped, white, 8-15 mm (0.3-0.6 in) long, 10-20 mm (0.4-0.8 in) in diameter. Fruits obovoid, white. Distribution: Bolivia, Paraguay, Uruguay, southern Brazil, and Argentina.

Rhipsalis clavata F. A. C. Weber 1892 Hatiora clavata (F. A. C. Weber) Moran 1953

Plants at first erect, becoming pendent, much branched, to 1 m (3.3 ft) or more long, basal portions determinate in growth but sometimes producing extension shoots of indeterminate growth. Stem segments not dimorphic, cylindrical to club shaped, truncate, to 5 cm (2 in) long, 2-3 mm in diameter, green or purplish, apically giving rise to whorls of two to seven younger segments. Areoles near the stem tips only. Flowers borne near or at the stem tips, bell shaped, white, to 15 mm (0.6 in) long. Fruits globose, whitish to pale pink. Distribution: Rio de Janeiro and Sao Paulo, Brazil.

Rhipsalis crispata (Haworth) Pfeiffer 1837 Epiphyllum chspatum Haworth 1830, Hariota crispata (Haworth) Lemaire 1839

Plants epiphytic, with many branches, more or less pendent, growth indeterminate. Stem segments leaflike, flat or sometimes three-winged, oblong, elliptical, or obovate, pale green, with notched to lobed margins, usually truncate basally, 6-10 cm (2.4-3.9 in) long, 2-4 cm (0.8-1.6 in) wide. Areoles minute, without bristles. Flowers borne singly or in clusters of two to four during the dry season, disk shaped, creamy white, 10-12 mm (0.4-0.5 in) in diameter. Fruits spherical, white. Distribution: Rio de Janeiro, Pernambuco, and Sao Paulo, Brazil.

Rhipsalis cuneata Britton & Rose 1923

Plants epiphytic shrubs, more or less pendulous, producing new segments from tips of old ones, growth indeterminate. Stem segments oblong to spatula shaped, wedge shaped basally, thin, green, deeply lobed to notched marginally, 8-12 cm (3.1-4.7 in) long, to 7 cm (2.8 in) broad. Areoles with one or two bristles. Flowers borne singly. Fruits globose, white. Distribution: Bolivia. Rhipsalis cuneata is poorly known.

Rhipsalis dissimilis (G. Lindberg) K. Schumann 1890 Lepismium dissimile G. Lindberg 1890 ?Rhipsalis chrysantha Loefgren 1915, ?Lepismiumchrysanthum (Loefgren) Backeberg 1959

Rhipsalis fíoccosa 615

?Rh¡psalis rígida Loefgren 1915, ?Lepismium rígidum (Loefgren) Backeberg 1959

Rhipsalis epiphyllanthoides Backeberg 1935, Lepismium epiphyllan-

thoides (Backeberg) Backeberg 1951 Lepismium marnieranum Backeberg 1963 Lepismium saxatile A. Friedrich & Redecker 1965, Rhipsalis saxatilis (A. Friedrich & Redecker) A. Friedrich & Redecker ex G. D. Rowley 1976

Rhipsalis spinescens J. A. Lombardi 1993

Plants epiphytic or lithophytic, loosely branching, growth determinate, erect at first, later arching, branching strictly acrotonic. Stem segments extremely variable, usually with 5-9 low ribs, sometimes three- to five-angled, other times nearly round in cross section, 5-14 cm (2-5.5 in) long, 4-10 mm (to 0.4 in) in diameter. Areoles with hairlike bristles 3-8 mm (to 0.3 in) long. Flowers borne laterally from sunken, woolly areoles, pale yellow, 10-15 mm (0.4-0.6 in) in diameter. Fruits nearly spherical, red with white bases. Distribution: Paraná and Sao Paulo, Brazil.

Rhipsalis elliptica G. Lindberg ex K. Schumann 1890 Rhipsalis chloroptera F. A. C. Weber 1898

Plants shrubby epiphytes, producing clumps of stems in clusters of three or four, pendent, growth indeterminate, 1-2 m (3.3-6.6 ft) long. Stem segments flat, broad, oblong to elliptical, separated by constrictions, margins faintly or strongly notched, dark green, 6-15 cm (2.4-5.9 in) long, 2.5-6 cm (1-2.4 in) broad. Areoles with some wool, sometimes with one bristle. Flowers borne laterally, one to five per areole, white, 8-9 mm (0.3-0.4 in) long, 12-20 mm (0.5-0.8 in) in diameter. Fruits globose to oblong, pink to red. Distribution: southern and southeastern Minas Gerais, Rio de Janeiro, Paraná, Santa Catarina, and Säo Paulo, Brazil.

Rhipsalis ewaldiana Barthlott & N. P. Taylor 1995

Plants epiphytic. Stem segments dimorphic; primary segments four-angled, growth indeterminate, to 60 cm (24 in) long, 4-5 mm in diameter, secondary segments mosdy three-angled, growth determinate, branching mesotonic, 3-6 cm (1.2-2.4 in) long. Flowers borne laterally on secondary segments, white, 14-20 mm (0.6-0.8 in) in diameter. Fruits pinkish, 8 mm (0.3 in) in diameter. Distribution: Rio de Janeiro, Brazil. The exact habitat of Rhipsalis ewaldiana is unclear.

Rhipsalis floccosa Salm-Dyck ex Pfeiffer 1837 Hariota floccosa (Salm-Dyck ex Pfeiffer) Lemaire 1839, Lepismium floccosum (Salm-Dyck ex Pfeiffer) Backeberg 1935 Rhipsalis pulvinigera G. Lindberg 1889, Lepismium pulvinigerum (G. Lindberg) Backeberg 1935, R. floccosa subsp.pulvinigera (G. Lindberg) Barthlott & N. P. Taylor 1995 Rhipsalis gibberula F. A. C. Weber 1892, Lepismium gibberulum (F. A.

C. Weber) Backeberg 1935 Rhipsalis tucumanensis F. A. C. Weber 1892, Hariota tucumanensls (F. A. C. Weber) Kuntze 1898, Lepismium tucumanense (F. A. C. Weber) Backeberg 1935, R. floccosa subsp. tucumanensis (F. A. C. Weber) Barthlott & N. P. Taylor 1995

Rhipsalis Elliptica

Rhipsalis dissimilis, photograph by Wilhelm Barthlott

Rhipsalis elliptica

Rhipsalis dissimilis, photograph by Wilhelm Barthlott

Rhipsalis elliptica

616 Rhipsalis floccosa

Rhipsalis pittieri Britton & Rose 1923, Lepismium pittieri (Britton & Rose) Backeberg 1959, R. floccosa subsp.pittieri (Britton & Rose) Barthlott & N.P.Taylor 1995 Rhipsalis flosculosa F. Ritter 1979

Rhipsalis hohenauensis F. Ritter 1979, R. floccosa subsp. hohenau-

ensis (F. Ritter) Barthlott & N. P. Taylor 1995 Rhipsalis monteazulensis F. Ritter 1979 Rhipsalis floccosa subsp. oreophila N. P. Taylor & Zappi 1998

Plants epiphytic or lithophytic, much branched, erect at first, becoming pendent, growth determinate, branching strictly acrotonic. Stem segments cylindrical, slender, round in cross section, often with raised podaria subtending the scale leaves, green but sometimes tinged with red or purple near the areoles, to 25 cm (9.8 in) long, 5-6 mm (0.2 in) in diameter. Areoles sunken, woolly, without bristles. Flowers borne laterally, greenish white to creamy white or golden yellow, 15-20 mm (0.6-0.8 in) in diameter, the buds rupturing the surface and surrounded by hairs. Fruits spherical, white, red, or pinkish, to 10 mm (0.4 in) in diameter. Distribution: Brazil, Venezuela, Paraguay, Argentina, Peru, and Bolivia. Rhipsalis floccosa is widely ranging and variable; six sub-

species are recognized. Subspecies floccosa has green stems, greenish white flowers, and white fruits; it occurs from northeastern to southeastern Brazil. Subspecies hohenauensis tends to have golden yellow flowers and white fruits; it occurs in eastern Paraguay, Argentina, and southern Brazil. Subspecies oreophila has almost perfectly cylindrical stem segments with no podaria, and very small flowers; it occurs in the highlands of central Bahia and northern Minas Gerais, Brazil. Subspecies pittieri tends to have more slender stem segments, and smaller flowers and fruits; it occurs in Venezuela. Subspecies pulvinigera has stem segments that are tinged red, larger flowers that may be more than 18 mm (0.7 in) in diameter, and pinkish fruits; it occurs in southeastern and southern Brazil. Subspecies tucumanensis is similar to subspecies pulvinigera but has stems without the red, and larger fruits that are red or white and to 10 mm (0.4 in) in diameter; it occurs in eastern Peru, Bolivia, and Argentina.

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