Mammillaria marksiana Krainz 1948

Plants solitary at first, later forming clumps. Stems flattened globose, light or yellowish green, 6-15 cm (2.4-5.9 in) high, 5-12 cm (2-4.7 in) in diameter. Tubercles pyramidal, weakly four-sided, axils woolly, sometimes densely so in the flowering zones. Spines difficult to distinguish as centrals and radi-als, variable in number, 4-21, thin, needle-like, golden yellow to brown, 5-8 mm (0.2-0.3 in) long. Flowers bright greenish yellow, to 15 mm (0.6 in) long and in diameter. Fruits club shaped, dark purplish red, to 20 mm (0.8 in) long. Seeds small, brown. Distribution: Durango and Sinaloa, Mexico.

Mammillaria mathildae Glass & R. Foster 1971 Mammillaria fittkauisubsp. mathildae (Glass and R. Foster) J. M. Luthy 1995

Plants forming clusters. Roots fibrous. Stems globose to flattened globose, 5-6 cm (2-2.4 in) high, to 5 cm (2 in) in diameter. Tubercles cylindrical or elongate rhomboid; some

Mammillaria marcosii
Mammillaria marksiana

axils with bristles. Central spines 4-5, slightly pubescent, reddish brown, 6-10 mm (0.2-0.4 in) long, one hooked, others straight. Radial spines 9-11, hairlike, white, slightly pubescent, 5-14 mm (0.2-0.6 in) long. Flowers pinkish white, to 20 mm (0.8 in) long. Fruits scarlet, 6-7 mm (0.2-0.3 in) long. Seeds brownish black. Distribution: Querétaro, Mexico.

Mammillaria matudae Bravo 1956 Mammillaria matudae var. serpentiformis Fittkau 1971 Mammillaria compacticaulis Reppenhagen 1987

Plants solitary to clustering basally. Stems elongate cylindrical, 10-20 cm (3.9-7.9 in) or more high, to 3 cm (1.2 in) in diameter. Tubercles conical, with dilute latex, axils naked. Central spine one, pointing upward, white, becoming brownish with age, needle-like, slightly flattened, to 5 mm long. Radial spines 18-20, translucent white with yellowish bases, 23 mm long, lower ones longest. Flowers funnelform, light reddish purple, to 12 mm (0.5 in) long. Fruits red with green tint, to 12 mm (0.5 in) long. Seeds light brown. Distribution: México, Mexico.

Mammillaria mazatlanensis K. Schumann ex Gürke 1901 Chilita mazatlanensis (K. Schumann ex Gürke) Orcutt 1926, Ebnerella mazatlanensis (K. Schumann ex Gürke) Buxbaum 1951 Neomammillaria occidentalis Britton & Rose 1923, Chilita occidentalis (Britton & Rose) Orcutt 1926, Mammillaria occidentalis (Britton & Rose) Boedeker 1933, Ebnerella occidentalis (Britton & Rose) Buxbaum 1951

Neomammillaria patonii Bravo 1931, Mammillaria patonii (Bravo) Backeberg 1931, M. mazatlanensis subsp.patonii (Bravo) D. R. Hunt 1998

Plants commonly forming clumps. Stems cylindrical, 12-15 cm (4.7-5.9 in) high, 4-5 cm (1.6-2 in) in diameter. Tubercles short, conical, without latex, axils naked or with one or two short bristles. Central spines 1-4, usually straight but

Mammillaria mathildae

sometimes hooked, reddish brown, upper ones in the same plane as the radials, to 15 mm (0.6 in) long. Radial spines 12-18, slender, white, needle-like, 5-10 mm (0.2-0.4 in) long. Flowers tubular, intensely carmine red to reddish purple to pink to purple, 30-40 mm (1.2-1.6 in) long. Fruits club shaped, brown, becoming reddish yellow, to 20 mm (0.8 in) long. Seeds black. Distribution: Colima, Sinaloa, Sonora, Jalisco, Michoacán, and Nayarit, Mexico.

Two subspecies of Mammillaria mazatlanensis are recognized. Subspecies mazatlanensis has 1-3 central spines, 12-18 radials, and pink to carmine red to reddish purple flowers; it occurs throughout the range of the species. Subspecies pa-tonii has 4 central spines, 13-15 radials, and purple flowers; it occurs on the Tres Marías Islands off the coast of Nayarit.

Mammillaria melaleuca Salm-Dyck 1850 Dolichothele melaleuca (Salm-Dyck) Boedeker 1933

Plants usually solitary, sometimes clustering with age. Stems globose, dark green, 6-7 cm (2.4-2.8 in) in diameter. Tubercles thick, ovate obtuse, robust, without latex, axils naked. Central spine usually one, purplish brown, slender, rigid. Radial spines 8-9, radiating, nearly straight, upper 4 a little longer, brown, lower ones white. Flowers yellow, to 30 mm (1.2 in) long. Fruits greenish to brownish. Seeds blackish brown. Distribution: Tamaulipas, Mexico.

Mammillaria melanocentra Poselger 1855 Mammillaria rubrograndis Reppenhagen & A. B. Lau 1979, M. melanocentra subsp. rubrograndis (Reppenhagen & A. B. Lau) D. R.Hunt 1997

Mammillaria linaresensis R. Wolf & F. Wolf 1990, M. melanocentra subsp. linaresensis (R. Wolf & F. Wolf) D. R. Hunt 1997

Plants solitary. Stems depressed globose, glaucous green, to 16 cm (6.3 in) high, 10-12 cm (3.9-4.7 in) in diameter. Tubercles firm, large, pyramidal, strongly four-angled, keeled, with latex, axils woolly at first, later naked. Central spine one, stout, awl shaped, black, usually ascending, to 25 mm (1 in) or more long. Radial spines 4-13, stout, unequal, white to black when young, later light gray, 6-22 mm (0.2-0.9 in) long, lowermost ones longest. Flowers whitish, bright deep pink, or bright carmine, to 20 mm (0.8 in) long and in diameter. Fruits club shaped, pink to scarlet, to 30 mm (1.2 in) long. Seeds brown. Distribution: Coahuila, Durango, Nuevo León, and Tamaulipas, Mexico.

Three subspecies of Mammillaria melanocentra are recognized. Subspecies melanocentra has 7-9 black to gray radial spines and bright, deep pink flowers; it occurs in Coahuila, Durango, and Nuevo León. Subspecies linaresensis has 4-6 white radial spines and whitish flowers; it occurs near

Linares, Nuevo Leon. Subspecies rubrograndis has 11-13 yellowish brown radial spines and bright carmine flowers; it occurs in Tamaulipas and Nuevo Leon.

Mammillaria mercadensis Patoni 1910 Neomammillaria mercadensis (Patoni) Britton & Rose 1923, Chilita mercadensis (Patoni) Orcutt 1926, Ebnerella mercadensis (Patoni) Buxbaum 1951 Mammillaria rettigiana Boedeker 1930, Ebnerella rettigiana (Boedeker) Buxbaum 1951, Chilita rettigiana (Boedeker) Buxbaum 1954

Mammillaria posseltiana Boedeker 1932, Ebnerella posseltiana (Boedeker) Buxbaum 1951, Chilita posseltiana (Boedeker) Buxbaum 1954

Mammillaria mazatlanensis subsp. mazatlanensis

Mammillaria melanocentra subsp. melanocentra

Mammillaria sinistrohamata Boedeker 1932, Ebnerella sinistrohamata (Boedeker) Buxbaum 1951, Chilita sinistrohamata (Boedeker) Bux-baum 1954

Mammillaria flavlhamata Backeberg 1962, not validly published Mammillariapennispinosa var. nazasensis Glass & R. Foster 1975, M. nazasensis (Glass & R. Foster) Reppenhagen 1987, M. pennispinosa subsp. nazasensis (Glass & R. Foster) D. R. Hunt 1997 Mammillaria berkiana A. B. Lau 1986

Plants solitary and clustering. Stems flattened globose to globose, olive green to dark green, to 8 cm (3.1 in) in diameter. Tubercles conical to short cylindrical, with latex often present at flowering time, axils without bristles, sparsely woolly. Central spines 1-7, awl shaped, yellow to deep red to reddish brown, with lighter bases, to 14mm (0.6 in) long,with one, occasionally two, hooked. Radial spines 13-35, white to yellow, variably finely pubescent, 6-9 mm (0.2-0.4 in) long. Flowers variable, bell shaped, more deep set, very pale magenta to magenta, 10-20 mm (0.4-0.8 in) long and in diameter. Fruits

Mammillaria microhelia usually club shaped, reddish green. Seeds brownish black, finely pitted. Distribution: Durango, Jalisco, Guanajuato, and Zacatecas, Mexico. Mammillaria mercadensis represents the main western phase of Mammillaria series Stylothelae. The Fitz Maurices (pers. comm.) believe there is insufficient character differentiation to recognize infraspecific taxa.

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