Lepismium brevispinum Barthlott 1987

Pfeiffera brevispina (Barthlott) P. V. Heath 1994, Rhipsalis brevispina

(Barthlott) Kimnach 1996 Acanthorhipsalis brevispina F. Ritter 1981, not validly published

Plants terrestrial, erect, 1-2 m (3.3-6.6 ft) high, branched. Stem segments green, leaflike, 2-3 cm (0.8-1.2 in) long, 2-4 cm (0.8-1.6 in) wide, 2-3 mm thick, with wavy margins. Areoles in the notches, with brownish wool, scaly. Spines 14, yellowish, 1-2 mm long. Flowers borne singly from the notches, white, to 4 mm long and 3 mm in diameter; pericarpels without bristles. Fruits globose, white, translucent. Distribution: Amazonas, Peru, at elevations of 1700-2000 m (5600-6600 ft).

Lepismium crenotum (Britton) Barthlott 1987 Hariota crenata Britton 1891, Acanthorhipsalis crenata (Britton) Britton & Rose 1923, Rhipsalis crenata (Britton) Vaupel 1926, Pfeiffera crenata (Britton) P. V. Heath 1994

Plants shrubby epiphytes, branching laterally. Stem segments narrowly oblong, sinuate, thin, to 30 cm (12 in) long and 6 cm (2.4 in) wide, with distinct midveins. Areoles large, with wool. Spines 3-8, to 4 mm long. Flowers borne laterally, small, red. Fruits berries to 7 mm (0.3 in) in diameter. Distribution: La Paz, Bolivia, at an elevation of 1800 m (5900 ft).

Lepismium cruciforme (Vellozo) Miquel 1838 Cactus cruciformis Vellozo 1825, conserved name; Hariota cruciformis (Vellozo) Kuntze 1891, Rhipsalis cruciformis (Vellozo) A. Castellanos 1925

Cereus myosurus Salm-Dyck 1827, Lepismium myosurus (Salm-Dyck)

Pfeiffer 1837, Rhipsalis myosurus (Salm-Dyck) Förster 1846 Lepismium commune Pfeiffer 1835

Lepismium knightii Pfeiffer 1835, Rhipsalis knightii( Pfeiffer) Förster

1846, Hariota knightii (Pfeiffer) Kuntze 1891 Lepismium tenue Pfeiffer 1835 Lepismium fluminense Miquel 1838 Rhipsalis mittlen Förster 1846

Lepismium ramosissimum Lemaire 1864, Rhipsalis ramosissima (Lemaire) K. Schumann 1890, Hariota ramosissimum (Lemaire) Kuntze 1891

Rhipsalis cavernosa G. Lindberg 1890, Lepismium cavernosum

(G. Lindberg) G. Lindberg ex K. Schumann 1890 Rhipsalis macropogon K. Schumann 1890

Rhipsalis squamulosa K. Schumann 1890, Hariota squamulosa

(K. Schumann) Kuntze 1891 Rhipsalis anceps F. A. C. Weber 1892, Lepismium anceps (F. A. C.

Weber) Borg 1937 Rhipsalis brevibarbis K. Schumann 1894 Rhipsalis radicans F. A. C. Weber 1896 Lepismium vollii Backeberg 1935

Plants lithophytic or epiphytic, semierect or creeping with some branching. Stem segments extremely variable, linear to lance shaped, three- to five-angled, winged, or flat, to 50 cm (20 in) long and 2 cm (0.8 in) wide, often with purple color, with distinctly notched margins. Areoles in the notches, with tufts of white wool. Spines absent or nearly so. Flowers borne laterally, one to five per areole, cream, yellow, pink, or rarely magenta, 1-1.3 cm (0.4-0.5 in) long. Fruits globose, purplish red, to 6 mm (0.2 in) in diameter. Distribution: occurring widely in northeastern Brazil, Paraguay, and Argentina. Lepismium cruciforme is variable and a number of varieties have been proposed.

Lepismium houlletianum (Lemaire) Barthlott 1987 Rhipsalis houlletiana Lemaire 1858, Hariota houiletiana (Lemaire)

Kuntze 1891, Acanthorhipsalis houlletiana (Lemaire) Volgin 1982 Rhipsalis regnellliG. Lindberg 1890 Rhipsalis houlletiii. D. Hooker 1894

Plants much branched, epiphytic, pendent shrubs to 2 m (6.6 ft) or more in length. Stem segments rounded at base, thin and flat above, becoming leaflike, 10-20 cm (3.9-7.9 in) long,

Rhipsalis Lepismium Cruciforme Plants

Lepismium aculeatum, photograph by Werner Rauh

Lepismium bolivianum, photograph by Urs Eggli

Lepismium cruciforme, photograph by Urs Eggli

Lepismium aculeatum, photograph by Werner Rauh

Lepismium bolivianum, photograph by Urs Eggli

Lepismium cruciforme, photograph by Urs Eggli

Lepismium houlletianum, photograph by Urs Eggli

Lepismium houlletianum, photograph by Urs Eggli

Pfeiffera Erecta
Lepismium ianthothele

1-5 cm (0.4-2 in) wide, pointed on both sides, deeply saw-toothed, vivid green, sometimes with reddish margins. Areoles naked. Flowers borne in clusters, funnelform to nearly bell shaped, not opening widely, creamy white, pendent, to 2 cm (0.8 in) long. Fruits spherical, red to black, 5-6 mm (0.2 in) in diameter. Distribution: widespread in eastern Brazil; also reported from Bolivia but not confirmed.

Lepismium ianthothele (Monville) Barthlott 1987 Cereus ianthothele Monv\\\el839, Pfeiffera ianthothele (Monville) F. A. C.

Weber 1898, Rhipsalis ianthothele (Monville) K. Brandegee 1902 ? Rhipsalis cereiformis Förster 1846

Pfeiffera erecta F. Ritter 1964, Lepismium erectum (F. Ritter) Siipplie 1990

Pfeiffera mataralensis F. Ritter 1964, Lepismium mataralense (F. Ritter) Süpplie 1990 Pfeiffera multigona F. Ritter 1964 Pfeiffera gracilis F. Ritter 1980 Pfeiffera ianthothele var. boliviana F. Ritter 1980 Pfeiffera ianthothele var. tarijensis F. Ritter 1980

Plants epiphytic and lithophytic shrubs with weak stems, spreading or pendent, moderately branched, 30-60 cm (1224 in) long. Stem segments usually four-angled, sometimes triangular,light green, 1.5-2 cm (0.6-0.8in) in diameter, with tuberculate margins. Areoles with six or seven bristles 4-5 mm long. Flowers borne laterally, sometimes terminally, white, yellowish, sometimes pinkish, 2-2.4 cm (0.8-0.9 in) long; pericarpels with bristles. Fruits spherical, violet-pink. Distribution: southeastern Bolivia and northwestern Argentina.

Lepismium incachacanum (Cárdenas) Barthlott 1987 Rhipsalis incachacana Cárdenas 1952, Acanthorhipsalis incachacana (Cárdenas) Volgin 1982

Plants epiphytic, pendent shrubs, freely branching, to 1.5 m (4.9 ft) long. Stem segments green, mostly flattened (two-ribbed), more or less wavy, 20-30 cm (7.9-12 in) long, 4-6 cm (1.6-2.4 in) wide, with notched margins. Areoles deeply sunken in notches, with wool and tufts of light-colored bristles and hairs. Flowers borne laterally, singly or in clusters of two, magenta, to 1.8 cm (0.7 in) long. Fruits five-angled, reddish. Distribution: Cochabamba, Bolivia, at an elevation of 2200 m (7200 ft).

Lepismium lorentzianum (Grisebach) Barthlott 1987 Rhipsalis lorentziana Grisebach 1879

Plants epiphytic, freely branching shrubs. Stem segments flattened or three-angled with wavy and notched margins, fresh green, 10-30 cm (3.9-12 in) long, 2-4 cm (0.8-1.6 in) wide, with prominent midveins. Areoles borne in notches, not deeply sunken, with little wool, without bristles or spines. Flowers borne singly, pendulous, small, bell shaped to fun-nelform, yellowish white; pericarpel four-angled, smooth. Fruits slightly four-angled, purple. Distribution: Andes of Bolivia and Argentina at elevations of 800-1200 m (26003900 ft).

Lepismium lumbricoides (Lemaire) Barthlott 1987 Cereus lumbricoides Lemaire 1839, Rhipsalis lumbricoides (Lemaire) Lemaire 1842

Rhipsalis sarmentacea Otto & Dietrich 1841, Hariota sarmentacea

(Otto & Dietrich) Kuntze 1891 Rhipsalis leucorhaphis K. Schumann 1900 Rhipsalis loefgrenii Britton & Rose 1923

Plants epiphytic shrubs, freely branching, clinging or creeping on the bark of trees and attached by aerial roots. Stem segments round, never winged, angled, or flattened, grayish to yellowish green, to 1 m (3.3 ft) or more long, 4-8 mm (to 0.3 in) in diameter. Areoles crowded, with short white wool, and five to eight stiff bristles 3-5 mm long, usually deciduous. Flowers borne laterally, rotate, white to yellowish, to 2.2 cm (0.9 in) long, 3-4 cm (1.2-1.6 in) in diameter; pericarpels naked. Fruits spherical, green, becoming purple. Distribution: widespread in Brazil, Uruguay, Argentina, and Bolivia, from sea level to 1900 m (6200 ft).

Lepismium micranthum (Vaupel) Barthlott 1987 Cereus micranthus Vaupel 1913, Acanthorhipsalis micrantha (Vaupel) Britton & Rose 1923, Lymanbensonia micrantha (Vaupel) Kimnach 1984, Pfeiffera micrantha (Vaupel) P. V. Heath 1994 Rhipsalis asperulaMaupel 1926

Plants lithophytic shrubs, erect at first, later sprawling to pendent, branching freely, to 1 m (3.3 ft) or more long. Stem segments two- or three-angled, lobed marginally, to 20 cm (7.9 in) long, 1.5-2 cm (0.6-0.8 in) in diameter. Areoles in notches, not deeply sunken, with dense yellowish wool. Spines as many as 6, some twisted and flattened. Flowers hummingbird pollinated, borne near the tips of older stems, broadly funnelform, purple, with the perianth parts only barely expanded; floral tubes well developed, to 1.2 cm (0.5 in) long; pericarpels with wool and a few bristles. Fruits spherical or oblong, more or less winged, with some wool and a few spines. Distribution: Puno, Peru, at an elevation of 2100 m (6900 ft).

Lepismium miyagawae (Barthlott 8c Rauh) Barthlott 1987 Pfeiffera miyagawae Barthlott & Rauh 1987, Rhipsalis miyagawae (Barthlott & Rauh) Kimnach 1996

Plants epiphytic shrubs, erect at first, later sprawling or pendent. Stem segments mostly three-angled, sometimes four-or five-angled, slightly tuberculate, flesh soft, to 1 m (3.3 ft)

Lepismium Lorentzianum

Lepismium lorentzianum, photograph by Werner Rauh

Lepismium lumbricoides, photograph by Urs Eggli

Lepismium incachacanum, photograph by Wilhelm Barthlott

Lepismium lorentzianum, photograph by Werner Rauh

Lepismium lumbricoides, photograph by Urs Eggli

Lepismium incachacanum, photograph by Wilhelm Barthlott long, 1.5-2 cm (0.6-0.8 in) in diameter. Areoles with short white wool. Spines 3-5, whitish, to 7 mm (0.3 in) long. Flowers borne near the stem tips, broadly bell shaped, brilliant orange, to 1.5 cm (0.6 in) long, 3-4 cm (1.2-1.6 in) in diameter; pericarpels tuberculate, with numerous bristles. Fruits spherical, whitish, translucent, to 1 cm (0.4 in) in diameter. Distribution: Santa Cruz, Bolivia, at an elevation of 600 m (2000 ft).

Lepismium monacanthum (Grisebach) Barthlott 1987 Rhipsalis monacantha Grisebach 1879, Acanthorhipsalis monacantha (Grisebach) Britton & Rose 1923, Pfeiffera monacantha (Grisebach) P. V. Heath 1994

Lepismium miyagawae, photograph by Wilhelm Barthlott

Acanthorhipsalis incahuasina Cárdenas 1954, Pfeiffera incahuasina

(Cárdenas) P. V. Heath 1994 Acanthorhipsalis monacantha var. samaipatana Cárdenas 1957, A. samaipatana (Cárdenas) F. Ritter 1981, Pfeiffera monacantha var. samaipatana (Cárdenas) P. V. Heath 1994

Plants epiphytic shrubs, freely branching, erect at first, later pendent. Stem segments flat to three-angled, with serrate margins, to 45 cm (18 in) long, 2-3 cm (0.8-1.2 in) wide. Areoles with yellowish wool. Spines 1-2, stout, sharp, black, 6-10 mm (0.2-0.4 in) long. Flowers borne laterally on upper segments, orange, to 1.5 cm (0.6 in) long; pericarpels naked. Fruits spherical, orange or pale pink. Distribution: southeastern Bolivia and northwestern Argentina at elevations of 300-2000 m(980-6600 ft).

Lepismium paranganiense (Cárdenas) Barthlott 1987 Acanthorhipsalis paranganiensis Cárdenas 1952, Rhipsalis paranganiensis (Cárdenas) Kimnach 1983, Pfeiffera paranganiensis (Cárdenas) P. V. Heath 1994

Plants lithophytic shrubs, densely branched, becoming pendent. Stem segments three- to five-angled, 2-4 m (6.6-13 ft) long, to 1 cm (0.4 in) in diameter, without aerial roots. Areoles with one to three short bristles. Spines 2-4, whitish, to 6 mm (0.2 in) long. Flowers borne in clusters at the stem tips, rotate, creamy white to yellowish, to 2 cm (0.8 in) long. Distribution: Cochabamba, Bolivia, at elevations of2600-3000 m (8500-9800 ft).

Brevispinum

Lepismium monacanthum

Lepismium paranganiense, photograph by Werner Rauh

Lepismium monacanthum

Brevispinum

Lepismium miyagawae, photograph by Wilhelm Barthlott

Lepismium paranganiense, photograph by Werner Rauh

Lepismium warmingianum (K. Schumann) Barthlott 1987

Rhipsalis warmingiana K. Schumann 1890 Rhipsalis linearis K. Schumann 1890, Lepismium lineare (K. Schumann) Barthlott 1991 Rhipsalis gonocarpa F. A. C. Weber 1892

Plants epiphytic shrubs, much branched, erect at first, then pendent. Stem segments slender, dark green with reddish edges, three- or four-angled or flat, to 30 cm (12 in) long, 0.7-2.5 cm (0.3-1 in) in diameter, with scalloped margins and blunt teeth. Areoles hairless and spineless, not deeply sunken. Flowers borne laterally, solitary, pendulous, bell shaped, white, 1-2 cm (0.4-0.8 in) long and in diameter; pericarpels angled. Fruits spherical, opaque, dark purple to black, 5-6 mm (0.2 in) in diameter. Distribution: Brazil, eastern Paraguay, and northeastern Argentina in seasonal forests at elevations to 1100 m (3600 ft).

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