Eriosyce rodentiophila f Ritter 1980

sandillön

Eriosyce megacarpa F. Ritter 1980, Rodentiophila megacarpa (F. Ritter) Y. I to 1981

Rodentiophila atacamensis F. Ritter ex Y. Itö 1981, not validly published

Plants subglobose to globose, 12-32 cm (4.7-13 in) in diameter, with stem tips covered by white wool. Roots fibrous, produced from small taproots. Ribs 18-35, obtuse, notched to form tubercles. Spines thick, needle-like, horn colored, becoming gray, curved upward. Central spines 4-8,30-50 mm

Eriosyce Megacarpa
Eriosyce odieri subsp. glabrescens

(1.2-2 in) long. Radial spines 12-15,20-30 mm (0.8-1.2 in) long. Flowers borne on older areoles, cream colored, 2.5-4 cm (1-1.6 in) long, 2.2-3 cm (0.9-1.2 in) in diameter; pericarpels and floral tubes covered by dense white wool and with stiff bristles. Fruits woolly, reddish yellow, 25-30 mm (1-1.2 in) long, not dehiscent. Distribution: along the coast from north of Taltal south to below Chanaral, Antofagasta, Chile.

Eriosyce senilis (Backeberg) Kattermann 1994 Neoporteria senilis Backeberg 1935 Neoporteria gerocephalaY. ltd 1957, Illegitimate name Neoporteria coimasensis F. Ritter 1963, Eriosyce senilis subsp. coimasensis (F. Ritter) Kattermann 1994 Neoporteria multicolor F. Ritter 1963 Neoporteria robusta F. Ritter 1963 Eriosyce senilis subsp. elquiensis Kattermann 1994

Plants subglobose to globose to elongate, 4-12 cm (1.6-4.7 in) in diameter. Taproots large, tuberous. Ribs 13-21, depressed or notched between the woolly areoles. Spines white to gray to yellow to brown, straight to strongly twisted, some hairlike, others stiff and needle-like to soft and flexible. Central spines 4-20,30-60 mm (1.2-2.4 in) long. Radial spines 15-40, 20-60 mm (0.8-2.4 in) long. Flowers borne on young areoles, narrow funnelform, various shades of car mine red, 1-3 cm (0.4-1.2 in) long and in diameter; pericarpels and floral tubes with tufts of wool and bristles. Fruits elongate, red, dehiscing by basal pores. Distribution: inland from Elqui Valley south to Cuesta Las Chilcas, Chile.

Three subspecies of Eriosyce senilis are recognized. Subspecies senilis has twisted spines and flowers with little wool and few bristles; it occurs primarily in the valley of the Rio Choapa. Subspecies coimasensis has straight spines and occurs some distance inland from Las Copimas south to Monte Negro. Subspecies elquiensis also has twisted spines but has flowers with dense wool and numerous bristles; it occurs throughout the central Elqui Valley.

Eriosyce sociabilis (F. Ritter) Kattermann 1994 Neoporteria sociabilis F. Ritter 1963

Plants globose to elongate, 3-10 cm (1.2-3.9 in) in diameter. Taproots short, conical. Ribs 13-16, notched and forming tubercles with sunken areoles. Spines irregular, mostly straight, dark brown to black. Central spines 6-12, 20-40 mm (0.8-1.6 in) long. Radial spines 6-20,3-20 mm (to 0.8

Eriosyce Sociabilis
Eriosyce senilis subsp. coimasensis
Eriosyce Sociabilis
Eriosyce rodentiophila
Neoporteria Flowers

Eriosyce sociabilis in) long. Flowers borne on young areoles, narrow funnel-form, fuchsia red, 2-3 cm (0.8-1.2 in) in diameter; peri-carpels and floral tubes with white bristles and wool. Fruits elongate, dehiscing by basal pores. Distribution: south of Totoral Bajo to south of Caldera, Atacama, Chile.

Eriosyce strausiana (K. Schumann) Kattermann 1994 Echinocactus strausianus K. Schumann 1901, Pyrrhocactus strau-sianus (K. Schumann) A. Berger 1929, Neoporteria strausiana (K. Schumann) Donald & G. D. Rowley 1966 Echinocactus catamarcensis Spegazzini 1905, Malacocarpus catamarcensis (Spegazzini) Britton & Rose 1922, Pyrrhocactus catamarcensis (Spegazzini) F. Ritter 1959 Echinocactus sanjuanensis Spegazzini 1905, Pyrrhocactus sanjuanen-sis (Spegazzini) Backeberg 1939, Neoporteria sanjuanensis (Spegazzini) Donald & G. D. Rowley 1966 Pyrrhocactus vollianus Backeberg 1957, Neoporteria volliana (Backeberg) Donald & G. D. Rowley 1966 Pyrrhocactus atrospinosus Backeberg 1963, not validly published Pyrrhocactus setiflorus Backeberg 1963, not validly published; Neoporteria setiflorus (Backeberg) Donald & G. D. Rowley 1966, not validly published

Neoporteria backebergii Donald &G. D. Rowley 1966, not validly published

Pyrrhocactus pachacoensis Rausch 1975, Eriosyce strausiana var.

pcichacoensis (Rausch) Kattermann 1994 Pyrrhocactus platyacanthus F. Ritter 1980

Plants globose to elongate, to 20 cm (7.9 in) high, 8-15 cm (3.1-5.9 in) in diameter, with naked stem tips. Roots fibrous, arising from short taproots. Ribs 12-14, obtuse, partly notched. Spines grayish black to dark horn colored, most curved upward. Central spines 1-8,30-40 mm (1.2-1.6 in) long. Radial spines 8-14,25-30 mm (1-1.2 in) long. Flowers borne on young areoles, broad fiinnelform, yellow, 3.5-4 cm (1.4-1.6 in) long, 3.5-5 cm (1.4-2 in) in diameter; peri-carpels and floral tubes with bristles and wool below. Fruits short oval, greenish, thin walled, to 7-10 mm (0.3-0.4 in), dehiscing by basal pores. Distribution: between San Juan and Mendoza, Argentina.

Two varieties of Eriosyce strausiana are recognized. Variety strausiana has stems to 20 cm (7.9 in) high and 8-15 cm (3.1-5.9 in) in diameter, 6-8 central spines, and flowers 4 cm ( 1.6 in) in diameter; it occurs near San José de Jachal. Variety pachacoensis has stems to 10 cm (3.9 in) high and to 8 cm (3.1 in) in diameter, 1-2 central spines, and flowers 3.5 cm (1.4 in) in diameter; it occurs near Pachaco.

Eriosyce subgibbosa (Haworth) Kattermann 1994

cact0 rosado, quisquito

Echinocactus subgibbosa Haworth 1831, Neoporteria subgibbosa (Haworth) Britton & Rose 1922

Echinocactus castaneoides Cels ex Salm-Dyck 1850, Neoporteria cas-

taneoldes (Cels ex Salm-Dyck) Werdermann 1938 Echinocactus rostratus Jacobi 1856, Chilenia rostrata (Jacobi) Backeberg 1939 Echinocactus cupulatus Förster 1861

Echinocactus clavatus Söhrens ex K. Schumann 1900, Neoporteria clavata (Söhrens ex K. Schumann) Werdermann 1939, Eriosyce subgibbosa subsp. clavata (Söhrens ex K. Schumann) Kattermann 1994

Chilenia subcylindrica Backeberg 1935, not validly published; Neoporteria subcylindrica (Backeberg) Backeberg 1951, not validly published

Neoporteria nigrihorrida Backeberg 1939, Eriosyce subgibbosa var.

nigrihorrida (Backeberg) Kattermann 1994 Neoporteria heteracantha W.T. Marshall 1941 Neoporteria litoralis F. Ritter 1959, Eriosyce subgibbosa var. litoralis

(F. Ritter) Kattermann 1994 Neoporteria castanea F. Ritter 1963, Eriosyce subgibbosa var. cas-

tanea (F. Ritter) Kattermann 1994 Neoporteria microsperma F. Ritter 1963 Neoporteria wagenknechtii F. Ritter 1963, Eriosyce subgibbosa var.

wagenknechtii (F. Ritter) Kattermann 1994 Neoporteria vallenarensis F. Ritter 1980, Eriosyce subgibbosa var. val-

lenarensis (F. Ritter) Kattermann 1994 Neoporteria ra pi fer a Y. Itö 1981

Plants globose to elongate, often becoming procumbent when large, to 100 cm (39 in) long, 6-25 cm (2.4-9.8 in) in diameter, yellowish to deep green, sometimes glaucous. Roots usually fibrous. Ribs 16-22, deeply notched between areoles, forming large tubercles. Spines yellow to brown or black, stiff, needle-like, mostly straight but sometimes strongly curved, not always readily distinguishable as centrals and radials. Central spines 1-16, thicker than the radials, to 40 mm (1.6 in) long. Radial spines 8-40, often thin, sometimes hairlike, 10-20 mm (0.4-0.8 in) long. Flowers borne on young areoles, sometimes more than one per areole, carmine red, variable in size; pericarpels and floral tubes with wool and sometimes bristles. Fruits elongate, bright red, dehiscing by basal pores. Distribution: along the coast from Huasco, Atacama, south to Concepción, Chile.

Eriosyce subgibbosa is variable and has received numerous names, many of which were placed in synonymy by Kattermann (1994), who recognized two subspecies, one with three varieties and the other with four, and who has provided additional details and distributions. Subspecies subgibbosa has yellowish green stems and comprises three varieties. Variety subgibbosa has elongate stems with 8-30 thin radial spines. Variety castanea has globose stems with thin radial spines. Variety litoralis has stems with 30-40 hairlike radial spines. Subspecies clavata has dark glaucous green stems and comprises four varieties. Variety clavata has slender stems with 11-13 thick radial spines. Variety nigrihorrida has mostly globose stems with dark radial spines. Variety vallenaren-sis has slender stems with 9-24 straight, thin radial spines, and flowers 3-5 cm (1.2-2 in) in diameter. Variety wagen-knechtii has very small flowers, less than 3 cm (1.2 in) in diameter.

Eriosyce taltalensis (Hutchison) Kattermann 1994 quisquito detaltal

Neoporteria taltalensis Hutchison 1955, Neochilenia taltalensis (Hutchison) Backeberg 1959, Pyrrhocactus taltalensis (Hutchison)

F. Ritter 1959

?Echinocactus fuscus Muehlenpfordt 1848, ?Neoporteria fusca (Muehlenpfordt) Britton & Rose 1922,?Chilenia fusca (Muehlenpfordt) Backeberg 1939, ?Neochilenia fusca (Muehlenpfordt) Backeberg 1942 ?Echinocactus hankeanus Förster 1885, ? Neochilenia hankeana (Förster) Dölz ex Backeberg 1942, ?Neoporteria hankeana (Förster) Donald &G.D. Rowley 1966 ?Echinocactus fobeanus Mieckley 1907, ?Chilenia fobeana (Mieckley) Backeberg 1939, ?Neochilenia fobeana (Mieckley) Backeberg 1942 Horridocactus paucicostatus F. Ritter 1959, Pyrrhocactus paucicosta-tus (F. Ritter) F. Ritter 1959, Neochilenia paucicostata (F. Ritter) Backeberg 1962, Neoporteria paucicostata (F. Ritter) Donald &

G. D. Rowley 1966, Eriosyce taltalensis subsp.paucicostata (F. Ritter) Kattermann 1994

Neochilenia eriocephala Backeberg 1959, not validly published; Neoporteria eriocephala (Backeberg) Donald & G. D. Rowley 1966, not validly published ?Neochilenia neofusca Backeberg 1959, not validly published Pyrrhocactus calderanus F. Ritter 1961, Neochilenia calderana (F. Ritter) Backeberg 1963, Neoporteria calderana (F. Ritter) Donald & G. D. Rowley 1966 Pyrrhocactus gracilis F. Ritter 1961, Neochilenia gracilis (F. Ritter)

Backeberg 1963 Neochilenia pulchella F. Ritter ex Backeberg 1962, not validly published; Pyrrhocactus pulchelius F. Ritter 1980, Neoporteria pulchella (F. Ritter) Ferryman 1991 Neoporteria woutersiana Backeberg 1962, not validly published Pyrrhocactus piiispinus F. Ritter 1962, Neochilenia pilispina (F. Ritter) Backeberg 1963, Neoporteria pilispina (F. Ritter) Donald & G. D. Rowley 1966, Eriosyce taltalensis subsp .pilispina (F. Ritter) Kattermann 1994

Pyrrhocactus scoparius F. Ritter 1962, Neochilenia scoparia (F. Ritter) Backeberg 1963, Neoporteria scoparia (F. Ritter) Donald & G. D. Rowley 1966

Pyrrhocactus echinus F. Ritter 1963, Horridocactus echinus (F. Ritter) Backeberg 1963, Neoporteria echinus (F. Ritter) Ferryman 1991, Eriosyce taltalensis subsp. echinus (F. Ritter) Kattermann 1994 Pyrrhocactus floccosus F. Ritter 1963, Neochilenia floccosa (F. Ritter) Backeberg 1963, Neoporteria floccosa (F. Ritter) J. Lode 1992, Eriosyce taltalensis var. floccosa (F. Ritter) Kattermann 1994 Pyrrhocactus glaucescens F. Ritter 1963, Neochilenia glaucescens

(F. Ritter) Backeberg 1963 Pyrrhocactus intermedius F. Ritter 1963, Neochilenia intermedia (F. Ritter) Backeberg 1963, Neoporteria intermedia (F. Ritter) Donald & G. D. Rowley 1966 Pyrrhocactus pygmaeus F. Ritter 1963, Neochilenia pygmaea (F. Ritter) Backeberg 1963, Eriosyce taltalensis var.pygmaea (F. Ritter) Kattermann 1994

Pyrrhocactus rupicola F. Ritter 1963, Neochilenia rupicola (F. Ritter) Backeberg 1963, Neoporteria rupicola (F. Ritter) Donald & G. D. Rowley 1966

Pyrrhocactus neohankeanus F. Ritter 1980, Neoporteria neohankeana

(F. Ritter) Ferryman 1991 Pyrrhocactus tenuis F. Ritter 1980

Pyrrhocactus transiens F. Ritter 1980, Neoporteria transiens (F. Ritter) Ferryman 1991

Eriosyce subgibbosa var. clavata

Pyrrhocactus Transiens

Eriosyce subgibbosa var. clavata

Eriosyce taltalensis subsp. paucicostata

Eriosyce taltalensis subsp. paucicostata

Plants subglobose to globose to elongate, 3-15 cm (1.2-5.9 in) in diameter. Taproots short, conical. Ribs 8-13, notched and forming tubercles with sunken areoles. Spines irregular, dark brown to black, straight to curved. Central spines 6-12, 20-40 mm (0.8-1.6 in) long. Radial spines 6-12,3-20 mm (to 0.8 in) long. Flowers borne on young areoles, narrow to broad funnelform, fuchsia red to pink to creamy white to pale yellow to white, 3-4 cm (1.2-1.6 in) long, 2-3 cm (0.81.2 in) in diameter; pericarpels and floral tubes with white wool and often twisted bristles. Fruits elongate, with seeds loose within, dehiscing by basal pores. Distribution: along the coast and into the nearby hills from Antofagasta south to Caldera, Atacama, Chile.

Eriosyce taltalensis is variable and has been subjected to abundant naming. Kattermann (1994) recognized four subspecies (two with two varieties each). Subspecies taltalensis has straight spines, red, pink, cream, or white flowers, and comprises two varieties. Variety taltalensis has red or cream flowers; variety pygmaea has white or pink flowers. Subspecies echinus has flowers white below, fuchsia red above, and comprises two varieties. Variety echinus has stem tips with short wool; variety floccosa has stem tips with long tufts of wool. Subspecies paucicostata has large, often elongate stems 6-15 cm (2.4-5.9 in) in diameter and yellow to red flowers. Subspecies pilispina has twisted spines and white flowers.

Eriosyce tenebrica (F. Ritter) Kattermann 1994 Thelocephala tenebrica F. Ritter 1980 Thelocephala fankhauseri F. Ritter 1980

Plants flattened to subglobose, blackish brown, 2-4 cm (0.81.6 in) in diameter. Taproots long, cone shaped, sometimes with necks. Ribs 12, forming into tubercles with areoles not

Eriosyce Tenebrica

sunken. Spines 3-6, all radials, thin, 1-2 mm long. Flowers borne on young areoles, funnelform, yellow to pale reddish brown, 3-5 cm (1.2-2 in) in diameter; pericarpels and floral tubes with long fluffy wool and many bristles. Fruits elongate, wind dispersed, dry, dehiscing by basal pores. Distribution: west of Domeyko to the valley of the Rio Choros, Chile.

Eriosyce umadeave (Werdermann) Kattermann 1994 Echlnocactus umadeave Werdermann 1931, Pyrrhocactus umadeave (Werdermann) Backeberg 1935, Neoporteria umadeave (Werdermann) Donald & G. D. Rowley 1966

Plants globose to somewhat elongate, dark green, to 40 cm (16 in) high, 10-20 cm (3.9-7.9 in) in diameter, with naked stem tips. Roots fibrous. Ribs 18-21, sometimes more, deeply notched, with elongate areoles. Spines 20-35 or more, to 45 mm (1.8 in) long, not easily recognized as centrals and radials, curved upward, white to violet. Flowers borne on older areoles, funnelform, pale yellow, 3-3.5 cm (1.2-1.4 in) in diameter; pericarpels and floral tubes with some wool and bristles. Fruits elongate, light brown, 3-4 cm (1.2-1.6 in) long. Distribution: Andes of western Salta, Argentina.

Eriosyce vertongenii (J. G. Lambert) D. R. Hunt 1997 Pyrrhocactus vertongenii}. G. Lambert 1995

Plants solitary, flattened to subglobose, greenish brown, to 3.5 cm (1.4 in) high and 5.5 cm (2.2 in) in diameter. Roots diffuse. Ribs 8-13, forming rounded tubercles with oval areoles. Central spine 0-1. Radial spines 5-9, stout, round in cross section, needle-like, more or less erect, slightly curved, with dark tips, to 15 mm (0.6 in) long. Flowers borne on young areoles, funnelform, light yellow to salmon, to 4 cm (1.6 in) long and in diameter; pericarpels and floral tubes with long fluffy wool. Fruits elongate, wind dispersed, dry, dehiscing by basal pores. Distribution: San Juan, Argentina.

Eriosyce villicumensis (Rausch) Kattermann 1994 Pyrrhocactus villicumensis Rausch 1974, Neoporteria villicumensis

(Rausch) Donald 1976 Pyrrhocactus melanacanthus Backeberg 1963, not validly published; Neoporteria melanacantha (Backeberg) Donald & G. D. Rowley 1966, not validly published

Plants subglobose, light gray-green with whitish surfaces, 49 cm (1.6-3.5 in) in diameter, with naked stem tips. Taproots short, conical. Ribs 7-11, broadly obtuse, barely notched between areoles. Spines black. Central spines 1-2, curved upward, 10-20 mm (0.4-0.8 in) long. Radial spines 3-4 pairs, flattened against the stem surfaces, 8-15 mm (0.3-0.6 in) long. Flowers borne on young areoles, funnelform, orange-yellow, 2.8-3.5 cm (1.1-1.4 in) long, to 3 cm (1.2 in) in diam

Escobaría 307

eter; pericarpels and floral tubes with white wool and brown bristles. Fruits globose, dark brown, dry, dehiscing by basal pores with lids. Distribution: Sierra Villicum, Argentina.

Eriosyce villosa (Monville) Kattermann 1994

quisco peludo

Cactus villosus Monville 1839, Neoporteria villosa (Monville) A. Berger

1929, Chileniopsis villosa (Monville) Backeberg 1935 Echlnocactus polyraphis Pfeiffer ex Salm-Dyck 1850, Chileniopsis polyraphis (Pfeiffer ex Salm-Dyck) Backeberg 1935, Neoporteria polyraphis (Pfeiffer ex Salm-Dyck) Backeberg 1951 Neoporteria atrispinosa Backeberg 1939 Neoporteria cephalophora Backeberg 1939 Neoporteria laniceps F. Ritter 1963

Plants subglobose to globose, becoming elongate, 4-10 cm (1.6-3.9 in) in diameter. Taproots large, tuberous. Ribs 1315, deeply notched between areoles. Spines mostly hairlike with some stiff and needle-like, brown to black. Central spines 4-12, usually stiffbut sometimes soft and flexible, 2028 mm (0.8-1.1 in) long. Radial spines 12-20,15-25 mm (0.6-1 in) long. Flowers borne on young areoles, carmine red, 2-2.3 cm (0.8-0.9 in) in diameter; pericarpels and floral tubes with long wool and glassy white bristles. Fruits elongate, red, dehiscing by basal pores. Distribution: Totoral Bajo south to Carrizallilo near Huasco, Atacama, Chile.

Eriosyce villosa

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