Echinopsis pseudomamillosa Crdenas 1959

Plants solitary, globose to broadly globose, to 8 cm (3.1 in) high and 12 cm (4.7 in) in diameter. Ribs about 18, notched, forming distinct tubercles. Areoles elliptical, cream colored, to 1.5 cm (0.6 in) apart. Spines 7-10, pectinate, very unequal, 4-7 pointing laterally, 2 pointing upward, one pointing downward, needle-like, gray with brown tips, 0.5-4 cm (0.21.6 in) long. Flowers borne laterally near the stem tips, open at night, funnelform, white, to 18 cm (7.1 in) long and 8 cm (3.1 in) in diameter. Distribution: Cercado province, Cochabamba, Bolivia.

Echinopsis peruviana subsp. peruviana

278 Echinopsis pugionacantha

Echinopsispugionacantha Rose & Boedeker 1931 Lobivia pugionacantha (Rose & Boedeker) Backeberg 1935 Echinopsis rossii Boedeker 1933, Lobivia rossii (Boedeker) Boedeker ex Backeberg & F. M. Knuth 1935, E. pugionacantha subsp. rossii (Boedeker) G. Navarro 1996 Echinopsis stoilenwerkiana Boedeker 1936 Lobivia culpinensis F. Ritter 1966 Lobivia cornuta Rausch 1972 Lobivia salitrensis Rausch 1974 Lobivia versicolor Rausch 1974 Lobivia adpressispina F. Ritter 1980 Lobivia campicola F. Ritter 1980

Plants usually solitary but sometimes clumping. Stems globose to ovoid, dull gray-green, 8-15 cm (3.1-5.9 in) high, to 4.5 cm (1.8 in) in diameter. Taproots long. Ribs about 17. Spines 4-7, reported as daggerlike though variable, directed sideways, pale yellow, to 2.5 cm (1 in) or more long. Flowers open during the day, fragrant, funnelform, yellow to pink to orange to red, to 4.5 cm (1.8 in) long and in diameter. Fruits globose, semidry, dehiscent. Distribution: southern Bolivia into northernmost Argentina.

Two subspecies of Echinopsis pugionacantha are recognized by some. Subspecies pugionacantha has reddish yellow flowers to 4.5 cm (1.8 in) long and in diameter; it occurs in both Argentina and Bolivia. Subspecies rossii has orange to reddish flowers; it occurs only in Bolivia.

Echinopsis quadratiumbonata (f. Ritter) d. r. Hunt 1997 Trichocereus quadratiumbonatus F. Ritter 1980

Plants shrubby, producing several erect branches basally, to 1 m (3.3 ft) high. Stems cylindrical, green, 4-5 cm (1.6-2 in) in diameter. Ribs 9-12 with transverse furrows. Areoles round, brown, about 1 cm (0.4 in) apart. Spines needle-like, white with brown tips, straight. Central spine usually one, stout, erect, 1-4 cm (0.4-1.6 in) long. Radial spines 7-10,0.4-1 cm (to 0.4 in) long. Flowers long, white, not opening widely. Fruits dark green, to 3.5 cm (1.4 in) long and in diameter. Distribution: Cordillera province, Santa Cruz, Bolivia.

Echinopsis rhodotricha K. Schumann 1900 ?Echinocactus forbesii Lehmann 1843,7 Echinopsis forbesii

(Lehmann) A. Dietrich 1849 Echinopsis chacoana Schütz 1949, E. rhodotricha subsp. chacoana (Schütz) P.J. Braun & Esteves 1995

Plants commonly forming clumps with erect or ascending branches. Stems cylindrical, dull gray-green, 30-80 cm (1231 in) high, to 9 cm (3.5 in) in diameter. Ribs 8-18, low, slightly wavy. Areoles 1.5-2.5 cm (0.6-1 in) apart. Spines yellowish with brown tips. Central spine 0-1, bent upward slightly, to 2.5 cm (1 in) long. Radial spines 4-8, spreading, slightly curved, to 2 cm (0.8 in) long. Flowers white, to 15 cm (5.9 in) long. Distribution: Paraguay and northern Argentina.

Two subspecies of Echinopsis rhodotricha are recognized. Subspecies rhodotricha has 8-13 ribs and 4-7 radial spines; it occurs in Paraguay and northeastern Argentina. Subspecies chacoana has 12-18 ribs and 7-8 radial spines; it occurs in the Chaco Boreal region of Paraguay.

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