Plants solitary, rarely forming branches. Stems depressed globose, dull dark green, 7-15 cm (2.8-5.9 in) high, 10-20 cm (3.9-7.9 in) in diameter. Ribs 11-13, prominent, acute. Areoles white to gray, 1.5-2 cm (0.6-0.8 in) apart. Spines horn colored to gray, with brown tips. Central spines 1-2. Radial spines 4-7. Flowers open at night, funnelform, white, sweet smelling, 18-20 cm (7.1-7.9 in) long, 10-13 cm (3.95.1 in) in diameter. Fruits globose, blackish green to brownish, very hairy, to 3 cm (1.2 in) long and 2.5 cm (1 in) in diameter. Distribution: between Asunción and Encarnación, southeastern Paraguay.
256 Echinopsis anástrophora
Echinopsis ancistrophora Spegazzini 1905 Pseudolobivia ancistrophora (Spegazzini) Backeberg 1941 Echinopsis polyancistra Backeberg 1933, Pseudolobivia polyancistra
(Backeberg) Backeberg 1942 Echinopsis hamatacantha Backeberg 1934, Pseudolobivia hamata-
cantha (Backeberg) Backeberg 1942 Echinopsis kratochviliana Backeberg 1934, Pseudolobivia kratochvil-
iana (Backeberg) Backeberg 1942 Echinopsis leucorhodantha Backeberg 1934, Pseudolobivia leucorho-
dantha (Backeberg) Backeberg ex Krainz 1942 Echinopsis pelecyrhachis Backeberg 1934, Pseudolobivia pelecyrha-
chis (Backeberg) Backeberg ex Krainz 1942 Echinopsis torreciilasensis Cárdenas 1956, Pseudolobivia torrecilla-
sensis (Cárdenas) Backeberg 1959 Lobivia arachnacantha Buining & F. Ritter 1956, Echinopsis arachna-cantha (Buining& F. Ritter) H. Friedrich 1974, E. ancistrophora subsp. arachnacantha (Buining & F. Ritter) Rausch 1977 Lobivia pojoensis Rausch 1968, Echinopsis ancistrophora subsp.
pojoensis (Rausch) Rausch 1977 Lobivia cardenasiana Rausch 1972, Echinopsis cardenasiana (Rausch) H. Friedrich 1974, E. ancistrophora subsp. cardenasiana (Rausch) Rausch 1977
Plants solitary or forming clumps. Stems depressed globose, glossy dark green, to 8 cm (3.1 in) in diameter. Ribs 15-20, straight, divided into many small tubercles. Central spine usually one, rarely as many as 4, sometimes absent, hooked, light brown, to 2 cm (0.8 in) long. Radial spines 3-10, flexible, spreading, curved backward, whitish, to 1.5 cm (0.6 in) long. Flowers borne laterally, open during the day, of all colors, from white, sometimes with pinkish tint, to deep red, unscented, 12-16 cm (4.7-6.3 in) long; floral tubes slender. Fruits oblong, green to greenish purple, somewhat dry, to 16 mm (0.6 in) long and 8 mm (0.3 in) in diameter. Distribution: western Argentina and southern Bolivia.
Four subspecies of Echinopsis ancistrophora are recognized. Subspecies ancistrophora has flowers of varying colors; it occurs from south of Salta, Argentina, north to just across
the border into southern Bolivia. Subspecies arachnacantha occurs farther north in the region of Samaipata, Santa Cruz, Bolivia, is smaller, and has yellow to yellow-orange flowers. Subspecies cardenasiana occurs in Tarija, Bolivia, and has violet-pink to red flowers. Subspecies pojoensis produces larger plants with reddish flowers; it occurs near Pojo, Cochabamba, Bolivia.
Echinopsis angelesii (R. Kiesling) G. D. Rowley 1980 Trichocereus angelesii R. Kiesling 1978
Plants shrubby, low growing, branching basally with many branches, forming low mats to 1 m (3.3 ft) high. Stems cylindrical, ascending, light green, opaque, 6-6.5 cm (2.4-2.6 in) in diameter. Ribs low, obtuse, notched. Areoles oval, with scanty wool. Spines rigid, awl shaped, expanded basally. Central spine one, to 2 cm (0.8 in) long. Radial spines 12, in lateral pairs, lowest one longer, to 1.5 cm (0.6 in). Flowers borne near the stem tips, bell shaped, white, to 20 cm (7.9 in) long and 14 cm (5.5 in) in diameter. Distribution: Guachipas department, Salta, Argentina.
Echinopsis antezanae (Cárdenas) H. Friedrich & G. D. Rowley 1974
Trichocereus antezanae Cárdenas 1953, Helianthocereus antezanae (Cárdenas) Backeberg 1955
Plants shrubby with one to five columnar branches arising basally, to 5 m (16 ft) high. Stems cylindrical, stout, to 25 cm (9.8 in) in diameter. Ribs 15-30. Areoles white, close set. Spines 10-27, thin, needle-like, flattened against the stem surface, often interlacing, some bent backward, yellowish to brownish, 2-10 cm (0.8-3.9 in) long. Flowers borne near the stem tips, open at night, white, 10-14 cm (3.9-5.5 in) long. Fruits globose, yellowish green, 2.5 cm (1 in) in diameter. Distribution: El Cerado province, Oruro, Bolivia.
Echinopsis arboricola (Kimnach) Mottram 1997 Trichocereus arboricola Kimnach 1990
Plants shrubby, epiphytic, erect at first, later pendent. Stems slender cylindrical, dark green, to 1.2 m (3.9 ft) or more long, 2.5-4 cm (1-1.6 in) in diameter, producing aerial roots. Ribs 9-11, compressed laterally, with nearly square or subconical tubercles to 3 mm high. Areoles on the tips of tubercles, with cream-colored wool, 3-4 mm apart. Spines 9-15, radiating, needle-like, sharp, yellowish to brownish, one longer than the others, 1-23 mm (to 0.9 in) long. Flowers borne near the stem tips, open at night, bell shaped, white, 12-13 cm (4.75.1 in) long, 13-15 cm (5.1-5.9 in) in diameter. Distribution: southern Bolivia and northern Argentina.
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