Cumulopuntia ignescens Vaupel F Ritter 1980

jala-jala, puskayo

Opuntia ignescens Vaupel 1913, Tephrocactus ignescens (Vaupel) Backeberg 1936

Plants forming dense cushions that are sometimes raised. Stem segments stout, broadly ovoid, tapered, somewhat pointed, prominently tuberculate apically, to 8 cm (3.1 in) long. Areoles about 15, absent from lower portion of segments, only those at the tips with spines. Spines as many as 20, erect, close set, some strong, others thinner, reddish brown, to 8 cm (3.1 in). Flowers red to orange-yellow, to 3.5 cm (1.4 in) long; pericarpels almost lacking areoles, bearing spines only at rims. Fruits barrel shaped, with many spines to 6 cm (2.4 in) long on the rims. Distribution: very high elevations on the western side of the Andes from Puno, Peru, south to Antofagasta, Chile, and into northern Bolivia.

Cumulopuntia mistiensis (Backeberg) E. F. Anderson 1999 Tephrocactus mistiensis Backeberg 1936, Opuntia mistiensis (Backeberg) G. D. Rowley 1958

Plants semiprostrate. Stem segments ovoid to elongate, slightly tuberculate when young, to 3 cm (1.2 in) long, olive green. Spines absent or rarely one, bristle-like, to 4 mm long. Dis tribution: Arequipa, Peru. Little is known about Cumulopuntia mistiensis.

Cumulopuntia pentlandii (Salm-Dyck) F. Ritter 1980 Opuntia pentlandii Salm-Dyck 1845, Pseudotephrocactus pentlandii (Salm-Dyck) Fric 1933, Tephrocactus pentlandii (Salm-Dyck) Backeberg 1936, Maihueniopsis pentlandii (Salm-Dyck) R. Kieslirig 1984

Tephrocactus subinermis Backeberg 1935, Opuntia subinermis

(Backeberg) Backeberg 1935 Tephrocactus rarissimus Backeberg 1936, Opuntia rarissima (Backeberg) Borg 1951

Tephrocactus wilkeanus Backeberg 1936, Opuntia wilkeana (Backeberg) Borg 1951

Plants forming more or less dense cushions. Stem segments small, globose to short ovoid, becoming broader than long with age, grass green, with 10-15 areoles on upper part, to 2.5 cm (1 in) long. Tubercles low, obscure, later absent. Areoles with yellowish wool, upper ones producing spines. Spines 16, borne on lower margins of areoles, whitish, bristle-like, bent away, diverging, sometimes absent from segments. Flowers short, yellow or red, to 3 cm (1.2 in) long; pericarpels broad, with a few spineless areoles above. Fruits barrel shaped, to 3.5 cm (1.4 in) long, nearly spineless. Distribution: highlands of Bolivia, Peru, and northern Argentina.

Cumulopuntia pyrrhacantha (K. Schumann) F. Ritter 1981

Opuntia pyrrhacantha K. Schumann 1899, Tephrocactus pyrrhacan-thus (K. Schumann) Backeberg 1958

Plants low growing, shrubby. Stem segments very small, ovoid to elliptical, tuberculate, shiny yellowish green, to 2 cm (0.8 in) long. Areoles covering segments, upper ones bearing spines. Spines 5-8, strong, yellowish red, to 4 cm (1.6 in) long. Flowers golden yellow, to 3.5 cm (1.4 in) long; pericar-

Cumulopuntia pentlandii, photograph by Urs Eggli

Cumulopuntia ignescens

Cumulopuntia pentlandii, photograph by Urs Eggli pels naked below, areoles above with glochids, spineless. Distribution: northern Chile into Peru.

Cumulopuntia rossiana (Heinrich & Backeberg) F. Ritter

1980

Tephrocactus pentlandiivar. rossianus Heinrich & Backeberg 1953,

Opuntia rossiana (Heinrich & Backeberg) D. R. Hunt 1997 Tephrocactus microclados Backeberg 1962, not validly published

Plants forming small, compact cushions. Stem segments to 2 cm (0.8 in) long, very short ovoid, gray-green, distinctly tu-berculate, with 10-15 areoles mostly above. Tubercles polygonal, sharply defined. Areoles 10-15, on upper sides of tubercles, narrowly elongate, white, bearing spines above. Spines 1-3, sometimes as many as 6, sometimes absent, unequal, strong, awl shaped, sometimes flattened, whitish to brownish, bent downward basally, to 1.5 cm (0.6 in) long. Flowers yellow, orange, brownish red, or red, to 4 cm ( 1.6 in) long; pericarpels obconical, with areoles bearing glochids near rims, sometimes with small spines. Fruits bucket shaped, wider than long, with long glochids. Distribution: Chuqui-saca, Bolivia, south to Jujuy, Argentina.

Cumulopuntia Sphaerica (Förster) E. F.Anderson 1999

choclo, corotilla, gaitito, perrito, puskaye Opuntia sphaerica Förster 1861, Tephrocactus sphaerlcus (Förster) Backeberg 1936

Cactus berteriColla 1834, rejected name; Echinocactus berteri (Colla) Gay 1847, Tephrocactus berteri (Colla) F. Ritter 1958, Cumulopuntia berteri (Colla) F. Ritter 1980, Opuntia berteri (Colla) A. E. Hoffmann 1989

Opuntia dimorpha Förster 1861, Tephrocactus dimorphus (Förster)

Backeberg 1942 Opuntia leucophaea Philippi 1891

Opuntia rauppiana K. Schumann 1899, Tephrocactus rauppianus (K. Schumann) Backeberg 1936, Cumulopuntia rauppiana (K. Schumann) F. Ritter 1981 Opuntia campestris Britton & Rose 1919, Tephrocactus campestris

(Britton & Rose) Backeberg 1935 Opuntia kuehnrichiana Werdermann & Backeberg 1931, Tephrocactus kuehnrichianus (Werdermann & Backeberg) Backeberg 1936, Cumulopuntia kuehnrichiana (Werdermann & Backeberg) F. Ritter

1981

Tephrocactus pseudorauppianus Backeberg 1936, Opuntia pseudo-

rauppiana (Backeberg) W.T. Marshall 1941 Tephrocactus mirus Rauh & Backeberg 1957, Opuntia mira (Rauh &

Backeberg) G. D. Rowley 1958 Tephrocactus muelierianus Backeberg 1957, Opuntia muelleriana

(Backeberg) G. D. Rowley 1958 Tephrocactus multiareolatus F. Ritter 1964, Cumulopuntia multiareo-

lata (F. Ritter) F. Ritter 1981 Cumulopuntia tubercularis F. Ritter 1980

Plants small, fragile, prostrate to semierect, moderately branched. Stem segments globose, 2.5-6 cm (1-2.4 in) long. Tubercles poorly developed or absent. Areoles 25-80 or more per segment, 3-6 mm (to 0.2 in) in diameter, close set, prominent, becoming contiguous at times, to bases of segments, most bearing spines. Spines 2-15, variable, stout, erect, straight or sometimes bent, unequal, reddish, becoming lighter with age, to 4 cm (1.6 in) long, occasionally reduced to a few short bristles. Flowers orange, to 4 cm (1.6 in) long; pericarpels broad, squat, with spines at or near the rims. Fruits globose, somewhat spiny. Distribution: widespread and variable, at lower elevations from Ayacucho, Peru, south into central Chile.

Cumulopuntia ticnamarensis F. Ritter 1980 Opuntia ticnamarensis (F. Ritter) D. R. Hunt 1997

Plants forming cushions. Stem segments subglobose, strongly tuberculate, to 4 cm (1.6 in) long. Tubercles sharply defined, polygonal. Areoles large, vertically elongate, 35-40, extending over the segment, most bearing spines. Spines 4-8, erect, straight, stout, brownish gray, to 5 cm (2 in) long, sometimes bristles. Flowers sulfur yellow, 4 cm (1.6 in) long, 5 cm (2 in) in diameter; pericarpels smooth, with blackish bristles on rims. Distribution: Tarapaca, Chile.

Cumulopuntia tortispina F. Ritter 1980 Opuntia guatinensis D. R. Hunt 1997

Plants forming cushions. Stem segments ellipsoidal to ovoid, to 7 cm (2.8 in) long, strongly tuberculate. Areoles large, sunken, not extending to segment bases, upper ones spinif-erous. Spines as many as 20, needle-like, spreading, sometimes curved outward from bases, brown, occasionally pectinate, to 5 cm (2 in) long. Flowers yellow, tinged with red or purple, to 5 cm (2 in) long; pericarpels with areoles only at or

Cumulopuntia sphaerica, photograph by Urs Eggli

near rims, with reddish brown bristles. Fruits cylindrical, smooth, to 4 cm (1.6 in) long, with bristles 3-5 mm long on rims. Distribution: Antofagasta, Chile.

Cumulopuntia tumida f. Ritter 1981 Opuntia tumida (F. Ritter) D. R. Hunt 1997

Plants laxly shrubby, small. Stem segments globose to short elliptical, to 10 cm (3.9 in) long. Areoles prominent when young, 90 or more, extending to segment bases. Spines 1-7, sometimes absent, unequal, the larger ones awl shaped, whitish with brown tips, to 3 cm (1.2 in) long. Flowers golden yellow, to 3.5 cm (1.4 in) long; pericarpels with areoles only above, bearing small fine spines. Fruits squat, to 3 cm (1.2 in) in diameter, with fine spines. Distribution: Arequipa, Peru.

Cumulopuntia unguispina (Backeberg) F. Ritter 1981 Opuntia unguispina Backeberg 1937, Tephrocactus unguispinus (Backeberg) Backeberg 1937

Plants small, fragile, prostrate to semierect, branching like a string of beads. Stem segments small, to 2.5 cm (1 in) long. Spines as many as 18, bent downward, the longer ones dark, to 1.5 cm (0.6 in) long. Flowers yellow, to 5 cm (2 in) in diameter. Distribution: Arequipa, Peru. Cumulopuntia unguispina is poorly known.

Cumulopuntia zehnderi (Rauh & Backeberg) F. Ritter 1981 Tephrocactus zehnderi Rauh & Backeberg 1957, Opuntia zehnderi

(Rauh & Backeberg) G. D. Rowley 1958 Tephrocactus aiboareolatus Backeberg 1963, Cumulopuntia alboareo-lata (Backeberg) F. Ritter 1981

Plants forming cushions to 50 cm (20 in) wide. Stem segments globose to elongate, tuberculate, with 20-25 areoles 3-4 cm (1.2-1.6 in) long. Tubercles nipple-like. Areoles to 1 cm (0.4 in) or more in diameter, sometimes elliptical, with abundant wool, extending to bases of segments. Spines 3-8, sometimes 10, on many areoles, unequal, spreading, sometimes curved downward, yellowish brown to whitish, to 3 cm (1.2 in) long. Flowers yellow, to 3 cm (1.2 in) long; pericarpels obconical, with 2-4 erect spines to 2.5 cm (1 in) long. Fruits spiny above, to 2 cm (0.8 in) long and 1.5 cm (0.6 in) in diameter. Distribution: Arequipa, Peru.

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