Armatocereus brevispinus J E Madsen 1989

Plants treelike, much branched, to 10 m (33 ft) high with well-defined trunks to 2 m (6.6 ft) high and to 50 cm (20 in) in diameter. Stem segments spreading, dark green with grayish tint, tapering at both ends, 40-80 cm (16-31 in) long, 7-12 cm (2.8-4.7 in) in diameter. Ribs 4-6, separated by deep sinuses, 2-4 cm (0.8-1.6 in) high. Spines 6-9, inconspicuous, light brown, becoming gray with age, spreading, 2-7 mm (to 0.3 in) long. Flowers borne horizontally, narrowly funnelform, white, 8 cm (3.1 in) long, 6 cm (2.4 in) in diameter. Fruits ovoid, green, 7-10 cm (2.8-3.9 in) long. Distribution: Catamayo to Loja, Loja, Ecuador.

Armatocereus cartwrightianus (Britton 8c Rose) Backeberg ex A. W. Hill 1938 cardon

Lemaireocereus cartwrightianus Britton & Rose 1920, Cereus cartwrightianus (Britton & Rose) Werdermann 1931

Plants shrubby or treelike, to 12 m (39 ft) high, usually with well-defined spiny trunks to 2 m (6.6 ft) high and 30 cm (12 in) in diameter. Stem segments dark green, tapering at both ends, 30-60 cm (12-24 in) long, 7-10 cm (2.8-3.9 in) in diameter. Ribs 6-9, separated by deep sinuses, 1.5-2.5 cm (0.6-1 in) high. Spines needle-like, brown, becoming grayish white with dark tips. Central spines 1-4, 2-10 cm (0.8-3.9 in) long. Radial spines 15-25, spreading, 0.5-2.5 cm (0.2-1 in) long. Flowers borne horizontally, white, 7-9 cm (2.8-3.5 in) long, 4-5 cm (1.6-2 in) in diameter. Fruits ovoid, green, becoming red, 5-9 cm (2-3.5 in) long. Distribution: southern Ecuador and northern Peru.

Armatocereus cartwrightianus

114 Armatocereus godingianus

Armatocereus godingianus (Britton & Rose) Backeberg ex

E. J. Salisbury 1947 Lemaireocereusgodingianus Britton & Rose 1920

Plants shrubby or treelike, to 10 m (33 ft) high with many spreading branches, often with smooth trunks to 1.5 m (4.9 ft) high and 15-50 cm (5.9-20 in) in diameter. Stem segments dark green, broadest below, 30-60 cm (12-24 in) long, 7-10 cm (2.8-3.9 in) in diameter. Ribs 7-11, separated by deep sinuses, 1.5-2.5 cm (0.6-1 in) high. Spines 15-25, brown or yellow, becoming gray, needle-like, flexible, spreading in all directions, 1-5 cm (0.4-2 in) long. Flowers borne horizontally, white, 7-9 cm (2.8-3.5 in) long, 5-7 cm (2-2.8 in) in diameter. Fruits ovoid, green, becoming brown, 6-13 cm (2.4-5.1 in) long. Distribution: along the Rio Chanchan, Ecuador.

Armatocereus humilis (Britton 8c Rose) Backeberg 1959 Lemaireocereus humilis Britton & Rose 1920, Ritterocereus humilis (Britton & Rose) Backeberg 1951

Plants shrubby, often forming dense thickets, branching infrequently. Stems weak, dark green, 1-4 m (3.3-13 ft) long, to 4 cm (1.6 in) in diameter. Ribs 3-6, interrupted, slightly undulate, with areoles in depressions. Spines 5-8, brown, becoming whitish, needle-like, 1-2 cm (0.4-0.8 in) long. Flowers spreading funnelform, greenish white, to 6 cm (2.4 in) long. Fruits globose, very spiny, to 4 cm (1.6 in) long. Distribution: Dagua Valley in the western cordillera of Colombia.

Armatocereus laetus (Kunth) Backeberg ex A. W. Hill 1938 Cactus laetus Kunth 1823, Lemaireocereus laetus (Kunth) Britton & Rose 1920

? Cereus ghiesbreghtii K. Schumann 1898, ? Armatocereus ghies-breghtii( K. Schumann) F. Ritter 1981

Plants treelike, much branched, 4-6 m (13-20 ft) high, branching freely near the ground from massive, short trunks. Stem segments columnar, gray-green, erect. Ribs 4-8, prominent. Spines 6-12, brown, becoming whitish or grayish with age, 1-2 cm (0.4-0.8 in) long. Flowers white, 7-8 cm (2.83.1 in) long, to 5 cm (2 in) in diameter. Fruits green, heavily spined. Distribution: northern Peru and possibly southern Ecuador. There is confusion regarding the nature of Armatocereus laetus; it may not be distinct from A. matucanensis.

Armatocereus mataranus F. Ritter 1966 Armatocereus mataranus var. ancashensis F. Ritter 1981, A. mataranus subsp. ancashensis (F. Ritter) Ostolaza 1998

Plants treelike, 3-7 m (9.8-23 ft) high, often with heavy branches. Stem segments bluish gray-green, 30-50 cm (12-

20 in) long, 10-15 cm (3.9-5.9 in) in diameter. Ribs 5-7, prominent, 4-5 cm (1.6-2 in) high. Spines awl-like, gray with darker tips. Central spines 3-8,1.5-6 cm (0.6-2.4 in) long. Radial spines 6-10,0.3-2 cm (to 0.8 in) long. Flowers white, to 9 cm (3.5 in) long. Fruits green, 6-8 cm (2.4-3.1 in) long. Distribution: Cajamarca and Ancash, Peru.

Armatocereus mataranus is poorly understood; it may be the same as A. laetus. Two subspecies of A. mataranus are recognized. Subspecies mataranus occurs in Cajamarca, whereas subspecies ancashensis is found only in Ancash.

Armatocereus laetus, photograph by Graham Charles

Armatocereus matucanensis Backeberg ex A. w. Hill 1938 Armatocereus arboreus Rauh & Backeberg 1957 Armatocereus churinensis Rauh & Backeberg 1957

Plants treelike, to 6 m (20 ft) high, branching mainly basally, usually without trunks. Stem segments erect, often clumping, gray-green with bluish tints, 30-50 cm (12-20 in) long,

7-13 cm (2.8-5.1 in) in diameter. Ribs 5-8, separated by shallow sinuses. Spines brown, becoming gray with dark tips, awl shaped, angular, flattened and slightly twisted. Central spines 1-4, 3-7 cm (1.2-2.8 in) long. Radial spines 8-14, spreading, 0.5-1.5 cm (0.2-0.6 in) long. Flowers borne horizontally to erect, narrowly funnelform, white, to 10 cm (3.9 in) long and 6 cm (2.4 in) in diameter. Fruits ovoid, green,

8-13 cm (3.1-5.1 in) long. Distribution: widespread throughout southern Ecuador and Peru.

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